As creatures like howler monkeys and toucans feast on that fruit, their droppings disperse seeds far and wide, planting new forest with every trip to the bathroom.
Like the beak in general in birds have many other functions, so they use it for display, nest construction, singing, thermoregulation in some cases like toucans or like hornbills ...
And the thing that was keeping them underground and keeping it from being released out into the atmosphere was that deep underground, underneath all the adorable sloths and toucans of Costa Rica, were chemolithoautotrophs.
As you close your eyes, your ears pick up hundreds of different sounds, from the cawing of toucans up above in the treetops to the rustling of hundreds of insects on the ground below.